Bothynus

Body length 16.2–30.8 mm, width 13.0–16.5 mm. Colour usually reddish to reddish brown, rarely dark brown or nearly entirely black with dark reddish tarsi. Head: clypeus subtrapezoidal to subtriangular, twice as wide as long or posterior width 2.7–3.2 times anterior width; dorsal surface rugopunctate, glabrous to moderately setose; clypeus surrounded by dense setae projecting from ventral surface. Frons transversely rugose or coarsely rugopunctate, usually with scarce setae on sides. Frontoclypeal region bearing a transverse, incomplete, raised carina not reaching lateral margins of head. Ocular canthus transverse, subrectangular, glabrous dorsally, ventrally bearing a transverse fringe of setae. Antennal club subequal in length to antennomeres 2–7 combined. Pronotum, in dorsal view, with a tubercle contiguous with middle of anterior margin; anterior area with a shallow to deep, rounded or oval cavity; surface transversely rugose to densely punctate; sides broadly rounded. Prosternal process columnar, cylindrical, projecting between coxae, apex oval. Scutellar plate mostly subtriangular, usually smooth. Elytra and part of pronotum may be covered with small setae. Legs: inner protarsal claw strongly curved, simple, equal in length to outer claw, or deeply incised or bearing a ventromedial process; protarsomere 5 subequal in length to protarsomeres 3–4 combined. Abdomen: tergite 7 with stridulatory apparatus formed by 1–2 bands of transverse striae; tergite 8 transverse, subtriangular, from rugose to densely punctate, glabrous to densely setose; sternite 8 from parabolic to subtriangular, with entire posterior margin. Aedeagus: phalobasis nearly 1.5 times longer than parameres; parameres, in caudal view, oval, long (1.5 times longer than wide), rounded laterally in basal half, apex usually with 2 oval lobes produced laterodorsally.

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